Emended diagnosis. Overall shape as well as gallery diameter and mesh-size varies but generally decreases towards the periphery of the trace. Branching points flattened parallel to the substrate surface and with rounded outline. Peripheral branches taper to pointed ends and have a jagged appearance. Galleries either developed as open channels and/or prostrate tunnels with few rhizoidal connections to the substrate surface.

Diagnosis:(Slightly modified after Wisshak2017,p. 43).‘Radiating, reticulate system of etched channelsor prostrate tubular borings at or immediately beneaththe surface of calcareous or bone substrates. Highdegree of anastomosis, leaving only a few peripheralbranches blind ending.’(See also H€antzschel1975,p.W127; Taylor and Wilson2003, table 2).